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Biology for Advanced Level Secondary Schools

           of it is produced during the light reaction.  photosynthesise,  even  if the  stomata  are
           The fact that the C  pathway produces  closed due to the presence of high levels
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           high  yields  under  high  concentration  of  of carbon dioxide  in the  bundle sheath
           carbon  dioxide  makes  photosynthesis  in   cells, about 20 to 120 times higher than
           C  plants more efficient in carboxylation   normal.
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           compared to C  plants, which under certain   Differences between C  and C  plants
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           conditions such as hot or dry conditions   The major differences between these
           undergo photorespiration.  The  PEP        two types of plants are  evident  in the
           carboxylase in the mesophyll of C  plants   photosynthesis  process. Generally, the
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           has a high affinity to carbon dioxide, and   basic metabolism of the C  and C  plants
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           it does not get competitively  inhibited   is similar, but the latter is more complex.
           by oxygen to cause photorespiration.       The C  plants are therefore evolutionarily
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           Although  carbon  dioxide  is a  limiting   more advanced than C  plants (Table 5.2).
           factor  in photosynthesis,  C  plants can                       3
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           Table 5.2: Differences between C  and C  plants
                                        3     4
                 C plants                                 C plants
                  3
                                                            4
             The first product in carbon dioxide fixation is   The first product in carbon dioxide
             a 3-carbon compound called phosphoglyceric   fixation is a 4-carbon compound called
             acid                                      oxaloacetic acid
             Photosynthesis is limited by low levels   Photosynthesis is not limited by low levels
             of atmospheric carbon dioxide, hence      of atmospheric carbon dioxide, hence
             photorespiration may occur under limited   photorespiration does not occur
             supply of carbon dioxide

             Carboxylation reaction is accomplished by   Carboxylation reaction is accomplished by
             the enzyme RuBP carboxylase to fix carbon   the enzymes RuBP carboxylase and PEP
             dioxide                                   carboxylase to fix carbon dioxide
             The carbondioxide acceptor is a 5-carbon   The carbondioxide acceptors are the
             compound called RuBP                      3-carbon compound (PEP) and the 5-carbon
                                                       compound (RuBP)
             Carbondioxide fixation occurs in bundle   Carbondioxide fixation occurs in both,
             sheath cells only                         mesophyll cells and bundle sheath cells
             They are less efficient in fixing carbon   They are more efficient in fixing carbon
             dioxide                                   dioxide
             They have only one type of chloroplast in   They have two types of cells, each with its
             their leaf bundle sheath cell             own type of chloroplast
             They do not tolerate hot and dry          They tolerate hot and dry conditions
             conditions. They are adapted to cool and
             wet environment
             Example of C  plants are soybeans spinach,   Example of C  plants are millet, sorghum and
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             rice and grasses                          sugarcane


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