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Gas exchange and respiration

           the respiratory surfaces. Thus, an efficient  a variety of respiratory surfaces, ranging
           respiratory surface must be moist for rapid  from gills in fish and tadpoles to lungs in
           exchange and transportation of respiratory  adult amphibians, reptiles, and mammals.
           gases.
                                                      Oxygen transport in vertebrates
           (g) Membrane permeability                  In  vertebrates  oxygen  is  transported  in
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           Since  the  membrane  form  the  boundary  two ways: as dissolved oxygen in blood
           between organism and its  extracellular  plasma and by means of red blood cells.
           environment,  it permeability  affects  The oxygen transported in solution form in
           the rate of gas exchange.  Therefore, a  the blood plasma accounts for only about
           respiratory surface must be permeable to  two percent (2%), and the remaining ninty
           allow gases to pass through.               eight percent (98%) is transported by the
                                                      red blood cells. The red blood cells have
           6.1.2 Oxygen and carbon dioxide            a red pigment called haemoglobin which
           transport in vertebrates                   is responsible for transport of oxygen and
           Oxygen and carbon dioxide are the  carbon dioxide in the blood. Structurally,
           respiratory gases that have to be transported  the  haemoglobin  molecule  consists of
           from one part of the body to another.  four Iron-containing parts and four protein
           Oxygen is transported from the respiratory   chains. Each haemoglobin molecule binds
           surfaces to the respiring body tissues and   to four oxygen molecules, forming  the
           carbon dioxide has to be transported from   oxyhaemoglobin molecule. This is carried
           the respiring body tissues to the respiratory   to individual cells in the body tissue where
           surfaces.  Vertebrates,  such as mammals,   it is released. The binding or combination
                                                      of oxygen and haemoglobin is a reversible
           birds, reptiles, amphibians and fish, have
                                                      process as shown in the equation below:






           According to the above equation, at high  the haemoglobin becomes. Usually the graph


           oxygen  concentration,  oxyhaemoglobin  is S-shaped or sigmoid.

           is formed,  whereas  at  low oxygen  The curve shows the following:

           concentration, oxyhaemoglobin dissociates  (a) At relatively low oxygen concentration,

           into haemoglobin and oxygen.                  there is uncombined haemoglobin in the
           This dissociation  releases oxygen from       blood and little or no oxyhaemoglobin
           haemoglobin. The balance can be shown         exists.  This is most likely to occur
           by an oxygen dissociation  curve of           in the body tissues, where oxygen


           oxyhaemoglobin  (Figure 6.4). In the          concentration is likely to be low.

           oxygen dissociation  curve,  the  greater   (b) At relatively high oxygen concentration,

           the  concentration  or partial  pressure of   haemoglobin combines with oxygen

           oxygen, the     more saturated with oxygen
                                                         in the blood to form oxyhaemoglobin.
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